Sub-human living beings - The living beings.
belonging to the realm of sub-human, which are other than the Aupapātika (3) (Deva (god) and Nāraka (infernal) and the human being and which always lead life in low (wretched) state.
aupapādikamanuṣyebhyaḥ śeṣāstiryagyonayaḥ.
(TaSū 4.28)
tirobhāvo nyagbliāgaḥ upabāhyatvamityarthaḥ tataḥ karmodayāpāditabhāvā tiryagyonirityā- khyāyate.(TaVā 4.27)